城市女孩安娜(艾米·亚当斯 Amy Adams 饰)与男友杰里米(亚当·斯科特 Adam Scott 饰)相恋四年,共同申请购买波士顿顶级豪华社区的高尚住宅。安娜本以为杰里米要向自己求婚,失望后听从父亲建议,飞往爱尔兰首都都柏林,打算依照爱尔兰传统,在闰年的2月29日这天向在都柏林参加医学会议的杰里米求婚。没想到飞机遭遇气流,迫降威尔士加的夫。安娜从加的夫雇船航行至爱尔兰丁格尔半岛,狼狈不堪中投宿当地青年德克兰(马修·古迪 Matthew Goode 饰)的海滨旅店。德克兰同意次日开车带安娜前往都柏林,一路上二人相互看不顺眼,争吵不休。争执中又遭遇车坏、暴雨、误点、假扮夫妻投宿农家等爆笑事件,可这一对冤家却也在不断的摩擦中彼此了解,情愫渐生。2月29日终于到来,安娜的一颗芳心究竟要托与何人呢?
At Christine's funeral, her foster son suddenly shows up, a grown-up man with whom the family has been out of contact for quite some time. He brought a letter, stating that Christine wants to be buried in Wentelen, to the bewilderment of her biological children, and even more so to that of her husband. In addition to that, all the relatives are surprised that the foster son of Christine apparently maintained a correspondence with Christine. Nevertheless, Christine's last wish is respected, and they decided to bury her in Wentelen, and wish to do so traditionally, meaning walking behind the hearse. During a walk of several days, all the relatives (foster sisters, foster father, alleged friends and real ones, the foster son, lovers) have enough time to get to know each other better. Because in the end they have no other conclusion than that nobody really knew anyone. Old intrigues are dug up, the foster sisters finally learn why the foster brother just disappeared from the family, and they can begin to develop a real image of the mother. Once Christine's final resting place is reached and she is buried, all family ties, good and bad, are shaken around.